Disclaimer: This is not an endorsement for or against the particular sites listed. They may or may not contain content that clashes with the site's rules. The sites are ordered alphabetically by URL (not counting the protocol or www).
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http://alanbishop.proboards.com/ (Home Grown Goodness forum)
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http://www.bigpumpkins.com/ (Discussion on such as giant pumpkins, other squash, watermelon, and muskmelons.)
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https://davesgarden.com/ (it's been around since the year 2000; one of the earliest gardening communities online; very useful for finding reviews of plant breeds and such; also, a nice place to trade seeds; paying members get to do more stuff)
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https://familycow.proboards.com/board/32/gardening (Keeping a Family Cow -> Gardening)
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https://garden.org/forums/view/eateat/•
https://www.gardenstew.com/ (GardenStew; an active community with general gardening forums.)
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http://gardening-forum.proboards.com/•
https://gardening.stackexchange.com/ (a Q/A site for home gardeners and landscapers)
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http://www.gardenplansireland.com/forum/ (IrishGardeners.com)
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https://www.gardenweb.com/ (It's been around
since at least 4 Jan 1996, and is the earliest gardening community I know about; it was acquired by Houzz in 2015; you have to enable private messaging for others to message you; please also edit your sidebar for a better experience, and don't expect it to work properly on a Kindle Fire HD 8 sixth edition)
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https://growingfruit.org/•
http://radishrain.321.s1.nabble.com/ (Radishrain: That's this very website.
See here for the site purpose.)
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https://www.helpfulgardener.com/ (Helpful Gardener; see the menu for the forums; it's been around since 2003)
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http://herb-gardening.freeforums.net/•
https://herb-talk.com/ (They have a forum for gardeners, but it's only part of the site.)
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http://louisianagardening.proboards.com/•
http://opensourceplantbreeding.org/forum/ (Open Source Plant Breeding Forum)
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http://perrenial-food-crops.proboards.com/ (Plant Breeding for Permaculture)
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https://plantvillage.psu.edu/ (a Q/A gardening site)
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http://seedsavingnetwork.proboards.com/•
http://notjusttomatoes.proboards.com/•
http://organicgroup.freeforums.net/ (The Organic Gardening Community)
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http://tatermater.proboards.com/ (Tom Wagner's forum)
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http://tatianastomatobase.com/wiki/Main_Page (Yeah, it's a wiki, but wikis have registered users, and a community feel.)
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http://thetomatogarden.proboards.com/•
https://tomaten-forum.com/ (Tomaten-Forum)
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https://www.tomatojunction.com/ (This surfaced just after an announcement of Tomatoville's end—note that it didn't end. Many Tomatoville members migrated, but other people joined, too, of course. The domain name changed to TomatoJunction.com on about 16 Dec 2019.)
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https://www.tomato-talk.com•
http://www.tomatoville.com/ (It's a great place for tomato information. Carolyn Male used the site a lot. The rules may be different than you expect, so be sure to pay attention to them. I've read that the original user-base largely migrated from Gardenweb.)
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http://vegetables.boards.net (The Savory Society: This is the predecessor to our forum! It's also a sister site, in case our forum goes down, or in case you want to use it.)
Retired communities:
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http://www.idigmygarden.com/ (It used to belong to Baker Creek Heirloom Seeds, but they sold it to the owner of Tomatoville, who eventually discontinued it.)
Climate:
BSk
USDA hardiness zone: 6